Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, 1024 Koajiro, Misaki, Miura, Kanagawa 238-0225, Japan

Kannonzaki Nature Museum, 4-1120 Kamoi, Yokosuka, Kanagawa 239-0813, Japan

Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, 1024 Koajiro, Misaki, Miura, Kanagawa 238-0225, Japan

Fisheries Research Institute, Japan Fisheries Research and Education Agency, 1-5939-22 Suido-cho, Chuo, Niigata 951-8121, Japan