Laboratory of Entomology, Obihiro University of Agriculture and Veterinary Medicine, 2-11 Inada-cho, Obihiro, Hokkaido 080-8555, Japan

Kanagawa Prefectural Museum of Natural History, 499 Iriuda, Odawara, Kanagawa 250-0031, Japan

Laboratory of Plant Ecology, Obihiro University of Agriculture and Veterinary Medicine, 2-11 Inada-cho, Obihiro, Hokkaido 080-8555, Japan
Faculty of Education, Hokkaido University of Education, Asahikawa Campus, 9 Hokumoncho, Asahikawa, Hokkaido 070-0825, Japan (present address)