Interdisciplinary Graduate School of Agriculture and Engineering, University of Miyazaki, 1-1 Gakuen-kibanadai-nishi, Miyazaki 889-2192, Japan
Nobeoka Marine Science Station, Center for Innovative Agriculture, University of Miyazaki, 376-6 Akamizu, Nobeoka, Miyazaki 889-0517, Japan

Unit of Marine Life Science, Faculty of Agriculture, University of Miyazaki, 1-1 Gakuen-kibanadai-nishi, Miyazaki 889-2192, Japan
Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250-0031, Japan

Nobeoka Marine Science Station, Center for Innovative Agriculture, University of Miyazaki, 376-6 Akamizu, Nobeoka, Miyazaki 889-0517, Japan
Unit of Marine Life Science, Faculty of Agriculture, University of Miyazaki, 1-1 Gakuen-kibanadai-nishi, Miyazaki 889-2192, Japan