This article has now been updated. Please use the final version.
https://orcid.org/0000-0001-8357-8593 Laboratory of Molecular Cell Dynamics, Faculty of Advanced Life Science, Hokkaido University PRIME, Japan Agency for Medical Research and Development
https://orcid.org/0000-0002-9328-5628 Department of Biological Sciences, Graduate School of Science, The University of Tokyo
Article ID: e190044
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