Graduate School of Engineering and Science, University of the Ryukyus

Rising Star Program, University of the Ryukyus
Marine Biodiversity Research Program, Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC)

Department of Natural History Sciences, Faculty of Science, Hokkaido University

Author's Organization：
Department of Chemistry, Biology and Marine Science, Faculty of Science, University of the Ryukyus