HEPATICAE OF BHUTAN II

Abstract

  Two species of Anthocerotae and 235 species of Hepaticae collected on two expeditions to Bhutan in 1979 and 1982 by D. G. Long are enumerated. The collection includes six species new to science which have been described elsewhere: Cololejeunea bhutanica, C. longiana, Bazzania bhutanica, Cephaloziella longii, Plagiochila monalata and P. parvivittata. Some 152 species are recorded from Bhutan for the first time, and in addition to the six new species, 37 are newly recorded for the Himalaya as a whole: Phaeoceros carolinianus, Temnoma setigerum, Bazzania assamica, B. revoluta, Cephalozia connivens, C. pleniceps, C. siamensis, Cephaloziella capillaris, C. crispata, C. herzogiana, C. kiaeri, Lophozia chichibuensis, L. diversiloba, Scaphophyllum speciosum, Marsupella boeckii, M. stoloniformis, Diplophyllum nanum, Lophocolea muricata, Plagiochila exigua, P. kitagawae, P. sawadae, P. trabeculata, P. uniformis, Plagiochilion fimbriatum, Schistochila macrodonta, Radula caduca, R. ceylanica, R. chinensis, Porella nitens, P. oblongifolia, P. subobtusa, Aphanolejeunea truncatifolia, Cheilolejeunea khasiana, Cololejeunea hasskarliana, C. madothecoides, C. obliqua and C. ocellata. Many of these represent significant extensions of known range, for example the discovery of Scaphophyllum, a genus previously considered endemic to Taiwan, is particularly noteworthy, as is the new discovery of the genera Aphanolejeunea, Schistochila and Temnoma in the Himalaya. The 44 additional species recorded previously from Bhutan are listed, bringing the total Anthocerotae and Hepaticae recorded to 281. Some ecological observations on the hepatic flora of the major vegetation zones of Bhutan are made together with an assessment of the major phytogeographic elements represented.