New species of fossil butterfly (Nymphalidae: Limenitidinae) from the Upper Pliocene to Lower Pleistocene Teragi Group, Hyogo Prefecture, Japan

Abstract

This study describes a new fossil butterfly species, Tacola kamitanii sp. nov., from the Upper Pliocene to the Lower Pleistocene Teragi Group in Hyogo Prefecture, Japan. The new fossil is characterized by remarkably large wings, with an estimated forewing length of 48 mm and a wingspan of 84 mm. This new fossil species belongs to the genus Tacola based on the following characteristics: both discal cells open, smoothly curved humeral veins, and a thick thorax and abdomen. However, it does not identify with any modern relatives of Tacola with a small discal cell, straight 1A+2A anal vein of the forewing, or long hindwing median vein. Therefore, the fossil was identified as a new species of the genus Tacola and one of the largest species in the subfamily Limenitidinae. The modern relatives of Tacola are distributed in the subtropics and tropics, while this fossil species may have survived in the temperate zone. This is the first named Limenitidini fossil and the youngest example of an extinct butterfly.

ZooBank registration: urn:lsid:zoobank.org:pub:FCF699C6-B28F-4916-91BE-B9C93A0BB0DC