Primate Research
Online ISSN : 1880-2117
Print ISSN : 0912-4047
ISSN-L : 0912-4047

This article has now been updated. Please use the final version.

Proceedings of the 33rd Annual Meeting of PSJ
JOURNAL FREE ACCESS Advance online publication

Article ID: 33.013

The final version of this article is now available: Vol. 33 (2017), No. 2 pp. 112-124
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