Author's Organization:Institute of Biological Control, Faculty of AGriculture, Kyushu University Institute of Biological Control, Faculty of AGriculture, Kyushu University:(Present address)Forest Products Research and Development Institute, College Institute of Biological Control, Faculty of AGriculture, Kyushu University:(Present address)Wuhan Institute of Virology, academia Sinica Institute of Biological Control, Faculty of Agriculture, Kyushu University