Japan College of Fish, 5-255-10 Misaki, Miura, Kanagawa 238-0243, Japan (present address)
Interdisciplinary Graduate School of Agriculture and Engineering, University of Miyazaki, 1-1 Gakuen-kibanadai-nishi, Miyazaki 889-2192, Japan
Nobeoka Marine Science Station, Field Science Center, University of Miyazaki, 376-6 Akamizu, Nobeoka, Miyazaki 889-0517, Japan

Hyuga City, Miyazaki, Japan

Hyuga City, Miyazaki, Japan

Nobeoka Marine Science Station, Field Science Center, University of Miyazaki, 376-6 Akamizu, Nobeoka, Miyazaki 889-0517, Japan
Department of Marine Biology and Environmental Sciences Faculty of Agriculture, University of Miyazaki, 1-1 Gakuen-kibanadai-nishi, Miyazaki 889-2192, Japan