STUDIES ON ANTIPODAL HEPATICAE. XII. GYMNOMITRIACEAE

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  Until about 1960 only three genera had been customarily assigned to the Gymnomitriaceae: Gymnomitrion Corda, Marsupella Dumort. and Prasanthus (G.) Lindb. & Arn. A fourth genus, Anomomarsupella Schust. (1969) proved to be identical to Eremonotus Kaal. ex Pears., which had been erroneously assigned to the Lophozioideae, following Müller (1939-40; 1951-58). New genera were assigned to the family during the 1960's: Herzogobryum (Grolle 1964), Acrolophozia (Schuster 1963, 1966), Poeltia Grolle (1967) based on taxa from the cold Antipodes or (Poeltia) Himalaya. In the present account, Nothogymnomitrion, Apomarsupella, Nanomarsupella, Lophonardia and Paramomitrion are described as new, all but Apomarsupella (3 spp.) are monotypic. In addition, another new monotypic genus, Cephalomitrion, is recognized for a paradoxical taxon assigned by Grolle to Herzogobryum, but almost surely more correctly assigned to the Cephaloziellaceae. Finally, the isolated genus Stephaniella Jack is tentatively assigned to the Gymnomitriaceae. Excluding Cephalomitrion, some 11 genera are thus recognized in Gymnomitriaceae, almost a four-fold increase in the number of taxa assigned to the family until the 1960's and a considerable increase from the 8 genera recognized in the key in Schuster (1974, pp. 9-11). With the increased number of genera assigned to the family, almost wholly in taxa from (a) the cold Antipodes; or (b) alpine areas of the Northern Andes, an attempt at arranging them in meaningful subfamilies becomes necessary. In Schuster (1984, p. 982) 3 subfamilies are recognized: Gymnomitrioideae, Eremonotoideae and Stephanielloideae. On succeeding pages these subfamilies and the Lophonardioideae are, albeit tentatively, accepted. All genera, and their perimeters, are discussed and/or described in some detail, and a conspectus of subfamilies is given. For several genera new species are described; for others, critical reassessments are given. Some 27 plates serve to illustrate critical details of all new genera and several taxa in previously described genera.