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Teruo TADOKORO
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Sonam WANGCHUK
Wildlife Conservation Division and CITES Management Authority, Department of Forests and Parks Service, Ministry of Agriculture and Forests
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Sonam WANGDI
Forest Division, Department of Forests and Parks Service, Ministry of Agriculture and Forests, Samtse, Bhutan
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Karma WANGDI
Ugyen Wangchuck Institute for Conservation, Ministry of Agriculture and Forests, Lamai Goempa, Bumthang, Bhutan
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Sherub
Ugyen Wangchuck Institute for Conservation, Ministry of Agriculture and Forests, Lamai Goempa, Bumthang, Bhutan
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Rinchen WANGDI
Research and Monitoring Section, Bumudeling Wildlife Sanctuary, Trashiyangtse, Bhutan
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Sangay DRUKUPA
Research and Monitoring Section, Bumudeling Wildlife Sanctuary, Trashiyangtse, Bhutan
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Motohiro HARADA
The University Museum, The University of Tokyo
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Motoki SAITO
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Toshiaki AOKI
The Research Institute of Evolutionary Biology
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Shuhei YAMAGUCHI
The Research Institute of Evolutionary Biology
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Yoshiko IGARASHI
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Yasuyuki WATANABE
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Masaya YAGO
The University Museum, The University of Tokyo