Abstract
In angiosperm, single pollen tube carrying sperm cells precisely targets to an ovule during fertilization processes. To ensure this one-on-one pairing, fertilized ovule inactivates the synergid cell to terminate the pollen tube attraction. We recently found a novel mechanism of inactivating the synergid cell by cell-fusion between the synergid cell and the endosperm. In this review, we describe the cell-fusion and other synergid inactivation mechanisms in Arabidopsis thaliana.