Hama Agricultural Regeneration Research Centre, Fukushima Agricultural Technology Centre, Minamisoma, Fukushima, 975-0036, Japan [Japan]

Hama Research Centre, Fukushima Agricultural Technology Centre, Soma, Fukushima, 979-2542, Japan [Japan]

The Faculty of Food and Agricultural Science, Fukushima University, Fukushima City, Fukushima, 960-1296, Japan [Japan]

Tokyo University of Agriculture and Technology, Fuchu, Tokyo, 183-8509, Japan [Japan]