Journal of the Japanese Society of Revegetation Technology
Online ISSN : 1884-3670
Print ISSN : 0916-7439
ISSN-L : 0916-7439
Volume 29, Issue 1
(2003 Aug.)
Displaying 1-50 of 65 articles from this issue
ORIGINAL ARTICLES
  • Satoshi OSAWA, Saori TOKUMARU, Takehiko KATSUNO
    2003 Volume 29 Issue 1 Pages 3-8
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The internal distribution of the vulnerable species Ludwigia peploidae ssp. stipulacea was investigated. The internal habitats of this species were recognized only twelve sites in 4 prefectures. On the characteristic of this species habitat, it was dominated by agricultural land use for paddy field cultivation, such as an abandoned paddy field, a ditch, a levee, a small reservoir, etc. The internal total aerial extent of Ludwigia peploidae ssp. stipulacea was 2,160 square maters. And 73% of its biomass was included in Chiba prefecture. Three habitats existed in a permanent water body, which was regarded as a location of a gentle disturbance. It was suggested that its community had been maintained by lengthening a leaf into the water body in order to avoid competition with other tall grass.
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  • Kimiyo ICHIKAWA, Hisayuki MAENAKA
    2003 Volume 29 Issue 1 Pages 9-14
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Potentilla discolor Bunge was grown in the research farm, and population dynamics of seedlings were investigated for three years. It was revealed that seedlings were found intensively within 60 cm around plants which produces mature seeds. The fact indicates that seeds dispersal distance were short. Many seedlings had died within one year after germination, whereas the survival rate of the seedlings which lived more than one year were high. Seedlings were often found in nearly bare areas than in high vegetation covered areas. These results suggest that ground management which can provide sparsely vegetation cover safe site is vital in conservation populations through regeneration from seedling.
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  • Kentaro MURAKAMI, Hisayuki MAENAKA, Yukihiro MORIMOTO
    2003 Volume 29 Issue 1 Pages 15-20
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    We investigated the number and coverage of homosporous pteridophyte species in 22 fragmented forest patches and mountain forests in Kyoto city area. Both the number and coverage of diploid species in the fragmented forest patches were less than those in mountain forests. In dry forest floors, polyploid apogamous species were dominant, because they had simpler reproduction systems that do not necessarily require liquid water to complete fertilization. As self-incompatible diploid species can only reproduce in moist habitats where spores are dispersed by other individuals, the immigrant ratio should be decreased according to the level of isolation of woodlots and the drying of the forest floor. The dominance ratio of polyploid apogamous species or forest floor diploid species can act as indices of urbanization.
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  • Kentaro YUTANI, Norio Tanaka, Takeshi TAKEMURA, Takashi ASAEDA
    2003 Volume 29 Issue 1 Pages 21-26
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    A experimental study was carried out to investigate the effects of shoots cutting time on the regrowth characteristics of Typha angustifolia. Field observation was conducted in the pond. Four 1 m by 1 m quadrats were demarcated in the T.angustifolia community, and surrounding area was kept undisturbed to use as a control area. In each quadrat, shoots were cut at about 20 cm above water surface in June, July, August and September 2002 and allowed to re-grow from cutting stubbles at all quadrats. The results of this study shows that June and July cut shoots were re-elongated up to the same height as the control. While delay of the cutting time showed decreased re-growth shoots biomass. Rhizome of August cut quadrat showed smallest biomass in December. Moreover, shoot cutting in August is found to be more suitable to suppress T.angustifolia expansion.
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  • Yusuke SIRONO, Norio TANAKA, Hajime WATANABE
    2003 Volume 29 Issue 1 Pages 27-32
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Long term dynamics of beachside vegetation, Carex kobomugi Ohwi.was evaluated by their growth model. The model was formulated by rhizome formation characteristics and long-term senescence with panicle formation obtained at Ryuyo beach. The model agrees well for the seasonal variation of their aboveground and belowground biomass. This model describes the future horizontal enlargement and distribution of their biomass. For horizontal distribution, we analyzed with 0.2, 0.4, 0.6, 0.8, 1.0 and 1.6 m transplanting distances. With 0.2 - 0.4 m distances, we couldn’t expect to control brown sand in wide area within a few years. Moreover, with 1.0 - 1.6 m distances, it took much time until controlling brown sand within expectable. The optimized transplant distances were found within 0.6 - 0.8 m by considering their enlargement and belowground biomass.
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  • Gan Zhu YAN, Kazuto SHIMA, Kyozo CHIBA
    2003 Volume 29 Issue 1 Pages 33-38
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The effect of mixed planting with grasses on growth of soil improving species under the nutrients poor condition was investigated. The amount of nodules and aboveground biomasses of the improving species decreased with increasing the number of mix planted grasses and increased with phosphorus fertilizer. The results suggest that a lack of phosphorus in soil was due to mix planting with grasses and it cause the decline of nodulation and low growth of soil improving species.
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  • Kanae ISHIMARU, Tetsuji IWAMA, Naoya OSAWA, Hiroshi TAKEDA
    2003 Volume 29 Issue 1 Pages 39-44
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Three broad-leaved species, Quercus serrata, Myrica rubra, and Quercus glauca, have been examined for four years to assess the influence of their initial size for crown development and biomass gain at Mt. Tanakami, Shiga Pref. Patterns of crown development and biomass gain according to the initial size were as follows; Q. serrata was most affected by the initial size, while M. rubra showed no significant difference in size. Quercus glauca indicated the same amount of growth regardless to the initial size. Thus, small individuals of Q. glauca and M. rubra, and large ones of Q. serrata may be suitable for planting in this area.
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  • Nobuo FUJIWARA, Yutaka YAMAGISHI, Takashi TANAKA, Keiji NIIJIMA, Keiko ...
    2003 Volume 29 Issue 1 Pages 45-50
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Measurement of the pruning brunch with maintenance of roadside trees and a questionnaire about pruning frequency to roadside trees administrators in all parts of Japan were carried out. And the influence of pruning upon the carbon fixation of roadside trees was considered, by means of comparing annual pruning brunch, which was calculated in various size of trees, with annual wood growth. As a result of the measurement, the dry-weight of the annual pruning brunch was increased with the growth of trees, and a dry-weight ratio of the annual pruning brunch to annual wood growth reached 45.3%, when the D.B.H. was 25cm, on an average of 4 species. And as a result of the questionnaire, it was known that a ratio of trees without pruning was 17.9% on an average of 6 species. These results showed that necessities to use the pruning brunch in effective and consider planting methods without pruning, as measures of the global warming in the future.
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  • Satoru TAKANASHI, Makoto TANI, Abdul Rahim NIK
    2003 Volume 29 Issue 1 Pages 51-55
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Surface conductance was studied using micrometeorological data obtained at a tropical rain forest in Peninsular Malaysia in 1998. Surface conductance could be estimated through the year using the surface conductance model expressed as the function of solar radiation and specific humidity deficit. Soil drought did not affect the characteristic of the surface conductance and transpiration rate in the driest period at this study site where soil and the air tend to dry in a tropical rain forest.
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  • Ryuichi TACHIBANA, Kanako NISHIMURA, Saori MIYAMOTO, Kenji FUKUNAGA
    2003 Volume 29 Issue 1 Pages 56-61
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Soil fauna is often indicated the characteristics of local environments. Species of the ants and the collembolas were identified in soil of the revegetated slopes and the neighbor forest. We discuss the relation among species, the vegetation structure and soil microbiota. As the vegetation structure of the revegetated slopes progressed, the species of the ants increased, and their composition became similar to that of the neighbor forest. The percentage of humus-feeding collembola increased and the composition of feeding groups became similar to that of the neighbor forest according to the decomposition of soil organic matter with the age of the slopes, involved with decreased Bacteria / Fungi ratio.
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  • Tatsuaki KOBAYASHI, Hirokuni NODA, Natsuko SUZUKI, Yosuke INADA, Yoshi ...
    2003 Volume 29 Issue 1 Pages 62-67
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The macrobenthic community in the main natural and man-made seashores in Tokyo Bay were surveyed in the summer of 2001. The result of the principle component analysis shows that the faunal composition depended on the principle habitat types as foreshore, estuary and lagoon. The difference between man-made and natural shores was not recognized. The result of multiple regression analysis of the main component shows that chemical oxygen demand and salinity of water and silt-clay content of bed-soil were effective variables. The fauna was however different between the bay-mouth and inner bay-area, influenced by regional condition. Crab community strongly depended on the development of marsh vegetation and the silt-clay content of bed-soil. The abundance of vegetation and topography in supra-tidal area supported the life of semi-terrestrial crabs.
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  • Yuri SUZUKI, Shozo SHIBATA, Kazuhiro TANAKA, Tetsuro SAKAI
    2003 Volume 29 Issue 1 Pages 68-73
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Using the image of high-resolution spacehome "IKONOS" we evaluated colors of identifiable tree leaves with high values in NDVI(Normalized Differential Vegetation Index)which is one of the indicator of vigorous trees by color model "HSV". Although it was difficult to discriminate species on the pixel scale, it was possible to classify species to some categories based on colors of leaves on the species scale. As a result, the possibility of more differences of colors between Cinnamomum camphora and Ligustrum lucidum, and C. camphora and Cedrus deodara.
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  • Taizo UCHIDA, Fuyuki TAZAKI, Junkoh MARUYAMA, Yohei SATO
    2003 Volume 29 Issue 1 Pages 74-79
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Phragmites australis(Cav.)Trin. and P. japonica Steud. communities were mowed at different heights in order to examine their regeneration response to above ground organs loss as one of the disturbance in waterfront(mowing at the ground level as high disturbance, mowing at 1/2 heights of plant height as low disturbance). Regardless of high and/or low disturbances, number of regenerated culms was approximately same in both species communities. However, it was indicated that P. australis and P. japonica suffered from more large stress further than low disturbance by high disturbance, because C/F ratio was higher and TNC content in rhizome was lower by high disturbance in both species communities. P. japonica communities, however, regenerated creeping culms vigor ously and seemed to form new-other communities by the culms under high disturbance.
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  • Hidehiro INAGAKI
    2003 Volume 29 Issue 1 Pages 80-84
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Some ecological characteristics of Lilium japonicum, particularly in respect to its life cycle, were investigated in native habitat. As the regrowth rate of foliage scale removing foliage part was only 15%, it is suggested that the present undergrowth management practices used to conserve the native habitat of mature lilies is preventing the growth of the young plants needed to replace older ones. Only 50% of the flowering-size bulbs displayed the following year. In addition, the survival rate of seed capsules, at three months after flowering, was greatly reduced because of weather conditions and insect damage. The percentage of repeat-flowering bulbs, which normally produced seeds, was estimated to be very low. Because L.japonicum is regarded as partially self-fertile as well as allogamous, it is considered that maintaining the population of pollinators is important for conservation of this lily in its native habitat
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  • Yoshiaki ISHII, Yu Ling LI, Qing Tu SI, Keiji SAKAMOTO, Lin He WANG, K ...
    2003 Volume 29 Issue 1 Pages 85-90
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    To clarify phenological aspect of adaptation of plants growing in arid region, seasonal changes in shoot elongation of four exotic(Artemisia ordosica, Caragana korshinskii, Salix psammophila, Sabina vulgaris)and one endemic(Salix matsudana)species were investigated for one growing season in Mu-Us desert, China. All species except A. ordosica started to grow in the beginning of May. Daily mean temperature in these days exceeded 10°C. Four native species stopped shoot elongation in July. But, S. matsudana continued to grow in summer season. A. ordosica and S. vulgaris on fixed sand dune restarted to grow just after rain. Only S.psammophila showed active growth near the water table.
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  • Tomoko KAMICHI, Tatsuaki KOBAYASHI, Masashi NOMURA
    2003 Volume 29 Issue 1 Pages 91-94
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The hybridization in the seeds which are used to restore the natural community of sect. Sciadorhodion was examined. Individuals with hybrid forms were observed in 27% of seedlings grown in the nursery. Interspesific compatibility in sect. Sciadorhodion reached 80% in the cross-test. In addition, Rhododendron dilatatum and R. kiyosumense shared the common pollinator in the garden. Therefore, the possibility that the hybrid exists in the nursery seedlings is high. It is not better to take seeds for restration from the garden in which the sect. Sciadorhodion is mixed.
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  • Makiko YAMAMOTO, Shigenobu TAMAI, Naoko TOKUCHI, Norikazu YAMANAKA
    2003 Volume 29 Issue 1 Pages 95-100
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Effects of fertilization on survival and growth were compared among four different Salix species including S. psammophila C.Wang et Ch. Y. Yang and S. matsudana Koidz., Chinese species for afforestation in semi-arid areas, and S.sieboldiana Blume and S. subfragilis Andersson, Japanese species. In this study, shoot growth, biomass and T/R rate were measured using cuttings of these species grown under different nutrient conditions. The S. psammophila and S. matsudana cuttings were more tolerant to heavy dressing of fertilizer than S.sieboldiana and S. subfragilis.
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  • Ken YOSHIKAWA, Naoko YASAKI, Keiji SAKAMOTO
    2003 Volume 29 Issue 1 Pages 101-106
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Leaves of three-year-old seedlings of Zelkova serrata and Quercus serrata were artificially defoliated in three levels, namely 0%, 50% and 100%. The changes in the phenology and photosynthetic activity were observed. One remarkable flash occurred just after the defoliation in both species. Leaf numbers of Z. serrata became the same level as before the treatment and these of Q. serrata exceeded the original numbers. Transpiration rate of both species increased after the defoliation treatment. Although the recovery of total leaf area of Z. serrata was restricted after the treatment, the productivity of Z. serrata could recover sufficiently because of high photosynthetic activity of old leaves flashed just before the defoliation treatment. The acceleration of photosynthetic activity of remained leaves of Q. serrata was enough to produce new leaves after the defoliation damage, and it showed almost the same level of productivity as the control condition.
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  • Yuling LI, Mariko HAYASHI, Keiji SAKAMOTO, Ken YOSHIKAWA
    2003 Volume 29 Issue 1 Pages 107-112
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Saplings of Sabia vulgaris Ant. and Juniperus chinensis var. kaizuka Hort. were grown under two levels of soil moisture condition, pF1.8 and pF4.2, and their top and root growth were measured. Drought stress prohibited the shoot elongation of S. vulgaris and promoted the growth of root system vertically. As a result, the T/R ratio of new growth decreased and the range of root system spread deeper as soil became dry. Both the top and the root of J. chinensis var. kaizuka were prohibited their growth under dry condition with the same T/R ratio of new growth as under wet condition. Most of the root system of J. chinensis var. kaizuka was distributed within 10 cm depth from the ground surface irrespective of the soil moisture conditions. The vertical elongation of root system without branching was the notable growth feature of S. vulgaris.
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  • Nozomi NAGAOKA, Norito OKADA, Takashi SHIMOMURA
    2003 Volume 29 Issue 1 Pages 113-118
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Using the video images, we investigated the characteristic of landscape evaluation(LE)structure of different planting form of the roof. The planting form was classified from the field study of the green roof, 12 cases(roof garden(RG)of 5 cases, planting of single species(PS)using turf(T)and Sedum(S), and mixed planting(MP)of 6 cases, non-planting of 1 case)were extracted, and the LE experiment was carried out using evaluation grid method and semantic differential method. In the profile analysis, the psychological evaluation(PE)of the PS(S)was the lowest of the 12 cases. Meanwhile, the PE of the PS(T)was higher than that of the RG. In the RG, PS, and MP, in the factor analysis, the evaluation factors(EF)of body sensation and nature cognition were extracted. In addition to these two kinds of EF, the EF of vision was extracted in the RG.
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  • Rie MATSUI, Kentaro MURAKAMI, Yukihiro MORIMOTO
    2003 Volume 29 Issue 1 Pages 119-124
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    We evaluated recovery of forest floor vegetation in the Expo ’70 Commemorative Park constructed about 30 years ago, using pteridophyte as an index. Comparing to urban woods near the park,the number of pteridophyte species reaches high level. Species-area curves of forest floors and artificial environments such as stone walls,ditches,watercourses were drawn,and we found that there are many pteridophyte species in deciduous forest and in artificial environments,like small watercourse under canopy.There is no significant differences in diversity index between forest floors and artificial environments,but composition of species is different. In the park,there are no species which prefer ridge environment,and species which prefer valley floor condition appear little.Existence of varied environments would be good for pteridophyte species to take root,and that will improve diversity of forest floor vegetation.
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  • Naoko MIKI, Bardel AL-BALUSHI, Yasuhito YAMAGUCHI, Ken YOSHIKAWA
    2003 Volume 29 Issue 1 Pages 125-130
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Structures of two mangrove forests of Avicennia marina(Forssk.)Vierh. under different site conditions on the coastal area of gulf of Oman in Arabian Peninsula were investigated in 2003. In Shinas, northern part of the Sultanate of Oman, the mangrove forest of A. marina was established along a channel from the sea. The maximum height(H)and the average crown projection area(CPA)of A. marina were about 6 m and 1.21 m2, respectively. A large number of seedlings were growing on the forest floor. On the other hand, H and CPA of A. marina in Quriyat, middle part of Oman, were about 4.5 m and 2.26 m2, respectively. The mangrove forest in Quriyat was surrounded on two sides by L shaped channel with a blocked mouth to the sea. No seedlings were growing in this land locked mangrove forest. The analysis of spatial distribution of individual trees by the m*-m relation suggested a random distribution of compact colonies with the size of 25 m2. Especially such colonial structure of spatial distribution was remarkable in the forest of Shinas with a large number of seedlings.
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  • Kimiko AZUMA, Tatsuaki KOBAYASHI
    2003 Volume 29 Issue 1 Pages 131-134
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The factors of spreading Pleioblastus chino Makino into secondary forests floor was studied. The analysis by quantification theory first family shows that the growth of P. chino was activated in soil pH more than 4.9 area, but the growth of P. chino was suppressed in soil pH under 4.3 area. The growth of P. chino was also activated in gentle inclined area of the bottom of slope and the border on path and coppicing area. The vigor of P. chino was less under the canopy of Pinus densiflora Sieb.. It is important that P. chino should be controlled in priority at areas where the growth of P. chino is activated.
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  • Field experiment for restoration of native Rhododendrons by seeding
    Junko MORIMOTO, Shozo SHIBATA, Shuzo HASEGAWA
    2003 Volume 29 Issue 1 Pages 135-140
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    We have tested several types of seedbeds for five years to build up the method for restoring the native Rhododendrons in secondary forests. Our findings are: 1)High germination rate is accomplished when the ground surface that seeds contact cannot erode easily and have high capacity of water content. 2)In addition to 1), certain amount of light intensity is required for seedling establishment. 3)Fertilized beds, which raise seedlings quickly to the juvenile size, enable the high survival rate of seedlings. We derived the effective techniques for the restoration of native Rhododendrons in a closed secondary forest. For the first place, it is necessary to increase the light intensity by cutting or thinning. Secondary, fertilized soils with rough particle size should be prepared for seedbeds. Seeds kept in the cool condition should be sown on the smooth seedbeds in early spring.
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  • Kimiyo ICHIKAWA, Katsuhiko INAMOTO, Motoaki DOI, Hideo IMANISHI
    2003 Volume 29 Issue 1 Pages 141-146
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Corms of crocus(Crocus medius Balb.)and bulbs of daffodil(Narcissus cyclamineus DC)and muscari(Muscari armeniacum Leithl. ex. Bak)were subjected to various temperatures for different periods before planting and planted in open field in fall. Exposure of crocus corms to 20°C stimulated their flower bud development and flowering, while exposure to 30°C delayed them. In daffodil and muscari, exposure of bulbs to 9°C filled their chilling requirement for inflorescence development resulting in early flowering. By combining some storage schemes, these geophytes exhibited ornamental value successively; from November to February in crocus and from January to April in daffodil and muscari. The longevity of flowers bloomed in winter was greatly improved as compared with that bloomed in fall and spring.
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  • Yusuke MORITA, Akihiro NAKAMURA
    2003 Volume 29 Issue 1 Pages 147-152
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Effects of flooding and following dry condition on growth and stomatal conductance were studied for 5 Salix species distributed in Kinki district. Adventitious roots were formed earlier in lower river distribution than that in mountainous and upper river distribution. Salix species distributed in lower river floodplain had high stomatal conductance in flooding condition, so these species seemed to have resistance for flooding. These species died in relative higher water content than in mountainous and upper river distribution in the following dry condition.
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  • Tomonori MITANI, Yoshiko KOSUGI, Makoto TANI, Satoru TAKANASHI, Tatsuy ...
    2003 Volume 29 Issue 1 Pages 153-158
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    Spatial and temporal variation of soil respiration rate was studied at an artificial forest of Hinoki(Chamaecyparis obtusa)at Kiryu experimental watershed in Shiga Prefecture, Japan. Soil respiration rate, soil temperature and soil water content have been measured on every 1 m grid in four 4m×4 m plots. Soil respiration rate has temporally depended on soil temperature. In each plot, the distribution of soil respiration rate has showed the frequency distribution centered on the mean of the plot. In the comparison of the plots, soil respiration rate was smaller at the water saturated point and also smaller at the droughty point influenced by water deficit in summer season. Annual soil respiration rate in each plot was estimated to be 454-566 gC/m2/yr by using the relationship between soil temperature and soil respiration rate.
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  • Junichi IMANISHI, Kayoko SUGIMOTO, Yukihiro MORIMOTO
    2003 Volume 29 Issue 1 Pages 159-164
    Published: 2003
    Released on J-STAGE: September 16, 2005
    JOURNAL FREE ACCESS
    The thermal response number(TRN)is the thermal inertial indicator that represents the degree of surface temperature change in response to input radiative energy. In this research, we conducted a canopy-level experiment using seedlings in order to demonstrate the possibility to detect vegetation water stress using TRN and the alternative thermal response number(ATRN), which we suggested as the alternative indicator that reduces the two observations of TRN into one using the air temperature about 30 min later from dawn. As a result, the TRN(dawn), which is calculated from the observed surface temperatures at around dawn and a time between 9:00 and 11:30 using airborne TABI equivalent thermal infrared sensor(relative precision of 0.1 °C)and the measured downward short- and long-wave radiation at ground, was potentially capable of detecting the existence of water stress of the vegetation about 4 in LAI.
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