(5) Suastus gremius gremius (Fabricius, 1798) The outbreak in the Yaeyama Isles in 1973 was the beginning of this hesperiid's northward expansion. It arrived in the Miyako Isles in 1974, in the Okinawa Isles in 1977, and finally arrived at Amami-Ooshima in 1998. After that, bypassing the record-free Isles of Tokara, Yakushima, Tanegashima, and Mishima, in 2006 it unexpectedly landed at the southern coast of the Satsuma Peninsula in Kyushu. Furthermore, in 2009 it crossed the Kagoshima Bay to the Oosumi Peninsula, and recently in 2010 this species was found on the Nichinan coasts, southeast of Miyazaki Prefecture (though reportedly on palms transported by a gardener). The sharp increase of this skipper's population in 1973 in the Yaeyama Isles indicates either that numerous migrants came from Taiwan, or that the population inceased following the founders' entry under some favorable weather conditions (e.g. mild winter). This species spreads by its own flight and/or human transportation of its foodplants with its early stages. Foodplants are 23 species of palms in 18 genera (Palmae). Most of them are introduced species, and only two are wild species which are also commonly planted. Accordingly human plantation of palms increased more habitable sites for this skipper. Wild species: Arenga tremula var. engleri, Livistoma chinensis var. subglobosa Introduce species: Archontophoenix alexandrae, Caryota urens, Cocos nucifera, Elaeis guineensis, Howeia belmoreana, Latania verschaffeltii, Mascarena verschaffeltii, M. lagenicaulis, Phoenix canariensis, P. reclinata, P. roebelenii, P. dactylifera, Ptychosperma macarthurii, P. elegans, Rhapis humilis, R. excelsa, Satakentia liukiuensis, Syagrus romoffia, Veichia merrillii, Washingtonia filifera, Trachycarpus fortunei * Although Chrysalidocarpus lutescens and Roystonea regia were recorded in the literature, I could neither find its early stages nor rear them on these palms. (6) Euploea mulciber barsine Fruhstorfer, 1904 Although this danaid was the latest among the butterflies which have shown northward expansion, it spread quickly from 1992 to 1999. In the Yaeyama Isles this species had been already recorded in 1952. Since then sporadic collection records followed in sequence, and records in the 1980's suggesting temporaly occurrences increased. Although its settlement was not confirmed on Okinawa-jima, collections were recorded every year from 1982 to 1984. The records on Okierabu-jima in 1981, on Amami-Ooshima in 1983, and in Tsushima (Nagasaki Pref.) in 1981 suggest that some emerging on Okinawa-jima moved northward. This species suddenly increased its number in 1992 in the Yaeyama Isles and Okinawa-jima, in 1995 in the Miyako Isles. Outbreaks in 1992 induced large scale northward expansion from 1995 to 1999 to Amami-Ooshima, the northernmost island of settlement as of 2010. Since 2000, this species has begun to reach further north, e.g. the Satsuma Peninsula and other localities in Kyushu. Its abundance in 1992 in the Yaeyama Isles was probably either due to many arrivals from Taiwan or due to the mild winter and high temperature in the following spring repeated for several years in succession, although no evidence is available. On Okinawa-jima this species was abundant as the mild winters continued from 1998 to 2001; on the other hand, in 2003 they declined sharply following the low temperature in the winter (the mean temperature of January 2003 was 15.7℃). In Taiwan this species exhibits the northward migration in the spring from its overwintering sites in the south. In the Nansei Isles it likely moves northward on favorable airflows. Its larval foodplants are trees and vines of the Moraceae and Apocynaceae (including Asclepiadaceae). Although there are several planted species, the situations of its foodplants have not been changed during its northward expansion. Moraceae: Ficus microcarpa, F. variegata, F.
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