Macrochelid Mites (Arachnida : Acari : Macrochelidae : Glyptholaspis, Macrocheles, Neopodocinum) Associated with Dung Beetles in Bali, Indonesia

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Ten species of mite of the family Macrochelidae (genera Glyptholaspis, Macrocheles, and Neopodocinum) are recorded from the body surface of dung beetles from Bali, Indonesia. Of those mites, two species, Macrocheles baliensis sp. nov. and M. monticola sp. nov., are described as new to science. Neopodocinum spinirostris (Berlese, 1910) is redescribed on the basis of adults and immature stages collected from Bali. Seven species, Glyptholaspis fimicola (Sellnick, 1931), Macrocheles hallidayi Walter and Krantz, 1986, M. krantzi Evans and Hyatt, 1963, M. merdarius (Berlese, 1889), M. oigru Walter and Krantz, 1986, Neopodocinum jaspersi (Oudemans, 1900), and N. spinirostris (Berlese, 1910), were recorded from Bali for the first time. Macrocheles arabaesc nom. nov. is proposed as a replacement name for M. scarabae Chinniah and Mohanasundaram, 1995 (preoccupied by M. scarabae Vishnupriya and Mohanasundaram, 1988).