The light-photosynthesis curve of the seedlings of two species of
Betura, Quercus, Acer, and two kinds of
Fraxinus grown under three light levels was compared in relation to their specific leaf area (
SLA) . With deeper shade, the
SLA showed a increase, the largest in both
Fraxinus, and the smallest in two kinds of
Betula. The half-saturation of photosynthesis of all tree species occurred in a low photosynthetic photon flux density (PPFD), with deeper shade, especially in
Quercus. With a reduction of PPFD, the initial slope of the light-photosynthesis curves of two species of
Betula showed a slight steepness, whereas that of the other tree species showed a great steepness. The dark respiration rate of leaves of all tree species decreased with a deepening of shade. The photosynthetic acclimation of both
Fraxinus was the greatest; this may be advantageous to surviving under a gap created in forest canopy. Two species of
Betula had the smallest photosynthetic acclimation to shade, and this characteristic may imply that they can not survive under shaded conditions. The acclimation to shade of
Quercus and
Acer was observed not in the
SLA, but in the photosynthesis; this might be an advantage for conserving photosynthates for expanding leaf area in shaded environments.
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