Here we provide new evidence suggesting that the North China is probably the locus of origin of the steppe mammoth, M. trogontherii, which had first adapted to the cool grassland environment in the mammoth lineage. The evidence comprises four teeth recently recovered from the lacustrine sediments of the Nihewan Formation at Majuangou site, Hebei Province. The sediments are dated earlier than 1.36Ma, which predates the first appearances of the steppe mammoth not only in Europe, but also in northeastern Siberia (ca.1.2-0.8Ma). Compared with all the contemporaneous mammoths, the molars of the four specimens have more plates (plate number: x17_1/2 on M_3 and x17x on M3), higher and narrower crown, thinner enamel, larger lamellar frequency and eruption angle. These characters can be regarded as advanced in the mammoth lineage, and the molars are barely indistinguishable from those of M. trogontherii from Europe not only in morphology, but also in all the measuring data. Thus they are referred to M. trogontherii herein. From chronological, distributional and morphological viewpoints, the specimens described here bridge the gaps between the earlier mammoth from China probably referable to M. meridionalis and the later M. trogontherii from the Europe and northeastern Siberia.
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