The moose deer, Alces alces, is the largest living deer which the main habitat of the species is the high-latitude coniferous forest of cool temperate to frigid zones of Eurasia and North America continents. The fossils of Alces alces were also known from 5 localities in Japan. Chronological ranges of 3 localities in Japan (Hanaizumi site, Kazaana-dokutsu site and Kumaishi-do cave) are limited from the end of MIS 3 to MIS 2. These Japanese records of Alces alces mark the southern limit of the species including both living and fossils. In 2008, a new fossil specimen of Alces alces, 17N III F18-2 was found from the late Late Pleistocene, T3 unit of Tategahana sand Member, Nojiri-ko Formation, Shinano Town, Nagano Prefecture, Central Japan. This is the first Alces alces record from the formation. The specimen is a fragmentary left dentary with almost complete M3. Two specimens, 16N III F18-36 (almost complete left lower M2) and 10N III E17-88 (lateral part of left dentary fragment), collected in previous excavations, are identified as the same individual as 17N III F18-2, since these three specimens fit precisely together. The age of the specimens is about 44,000y. B.P. (MIS3), based on tephrochronological and AMS^<14>C method dating data. This occurrence indicates sympatric distribution of two large-sized cervids, Sinomegaceros yabei and Alces alces. And these findings provide important knowledge on the environmental transition of Nojiri-ko and its surrounding area through Late Pleistocene. The new fossil Alces alces (17N III F18-2, 16N III F18-36 and 10N III E17-88) from the Nojiri-ko Formation marks the oldest record of the species from Japan and indicates that Alces alces had existed in Japan before MIS3. Based on the paleoclimatological and sea bottom topographical studies, it is supposed that the migration of Alces alces from Eurasia to Honshu, across Hokkaido, was in the cool period of MIS4.
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