The factors determining water-use efficiency, net assimilation rate and transpiration rate were investigated using two parameters α(ratio of total conductance to CO2 to total conductance to H2O) and β(ratio of total conductance to latent heat to total conductance to sensible heat, or surface moisture availability). The characteristics of water-use efficiency were detected from several data sets of field observations with porometric method as follows, (1) Water-use efficiency was higher in the morning than in the afternoon. (2) Water-use efficiency was higher in winter than in summer. (3) Water-use efficiency values were scattered mainly from 10-1 to 102 (mg/g), and the fluctuation of water-use efficiency with species was smaller than that with environmental conditions. These characteristics of water-use efficiency reflect the difference of the diurnal and seasonal variation patterns of transpiration rate and net assimilation rate. The phenomena (1) and (2) were mainly caused by the differences in vapor pressure deficit, and partly by the differnces in α, and the phenomenon (3) was caused by the little differences in α with species. Furthermore, the ranges of values of water-use efficiency, net assimilation rate and transpiration rate were discussed using two parameters, α and β.
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