This research concerns developmental processes in the use of stone tools by wild chimpanzees in Bossou, Guinea. Previous studies had shown that chimpanzees in Bossou began at age 3.5 years to use a pair of stones as a hammer and anvil to crack open nuts. A field experiment was conducted to clarify developmental processes, at an outdoor laboratory where stones and oil-palm nuts were provided by the experimenter. We directly observed and video-recorded infant chimpanzees' object manipulation in a social context, which is a prerequisite for nut-cracking. This paper is based on data collected between 1992 and 1994 consisting of all episodes of behavior related to nut/stone manipulation and nut-cracking. Six infant chimpanzees under the age of 3 years were observed. A total of 310 behavioral episodes were grouped first into 5 major behavioral categories: (1) manipulation of nuts, (2) manipulation of stones, (3) manipulation of both nuts and stones, (4) manipulation of nuts and/or stones while interacting with the other individuals, and (5) interaction with the other individuals who perform nut-cracking. Each of these categories was subdivided into 2-4 subcategories by looking at qualitative differences in behavior, such as direction of manipulation, plurality of manipulated objects, and temporal sequence. Relative frequencies of behavioral episodes for each category or subcategory was compared among three age groups: 0.5, 1.5, and 2.5 year olds. The development of manipulation skills may be characterized as follows. Infants increased in the number of behaviors in which they manipulated both nuts and stones successively and/or simultaneously, while interacting with the other individuals. These other individuals may play an important role as a safe-base for exploration of nut/stone manipulation, and as a model for emulation learning.
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