The Dicnemonaceae are haplolepideous mosses restricted to the southern Hemisphere and marked by a high degree (73% of its species) of endemism. Previously comprised of five genera, only two, Dicnemon and Eucamptodon, are retained in the family.
Dicnemon is confined to Austral-asia (New Guinea, Vanuatu, New Caledonia, Australia and New Zealand). It is comprised of five sections, thirteen species and one variety: sec. 1. Dicnemon - D. calycinum, D. dixonianum sp. nov., D. semicryptum; sec. 2. Synodonria stat. nov. - D. spathoideum, D. cuspidatum; sec. 3. Hypnopsis stat. nov. - D. pancheri var. pancheri, D. pancheri var. falcata stat. nov., D. connivens, D. seriatum comb. nov., sec. 4. Brauniella stat. nov. - D. subpiliferum comb. nov., D. planifolium, D. cochlearifolium comb. nov.; sec. 5. Fleischeria sec. nov. - D. robbinsii, comb. nov., D. novae-guinea comb. nov.
Eucamptodon occurs in Austral-asia and southern South America. It is reduced to two species and one variety: E. perichaetialis (Juan Fernandez Islands, Chile and southwestern Argentina); E. muelleri var. muelleri (eastern Australia, New Caledonia, and Norfolk Island); and E. muelleri var. norrisii var. nov. (Queensland, Australia).
The family is characterized by a derived haplolepideous peristome, massive precociously germinated, endosporically developed protonemata, specialized leaf nematogens that give rise to leaf rhizoids, long sheathing perichaetial leaves and the absence (variably expressed in some species) of a costa.
The Dicnemonaceae appear to be recently derived and closely related to an atypical element of the Dicranaceae (Braunfelsia, Parisia, and some members of Dicranoloma).
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