1. The glands, distributed in the narrow area contiguous to the proventriculus have been regarded as cardiac glands that secreted serum or mucus. However, they contain pepsinogen granules besides several kinds of mucosubstances. 2. Several simple tubular glands, distributed in the immediate vicinity of the proventriculus (A glands), have few parietal cells, and the glands strongly reacted to PAS and AB (pH 2.5), but not to AB (pH 0.5). 3. The A glands contain sialomucin in the glandular neck, weak acid and neutral mucopolysaccharides in the body and base of the glands, but no strong acid mucopolysaccharide. 4. The A glands contain fine pepsinogen granules in the glandular cells of the basal portion. 5. A great number of parietal cells appear in the glands contiguous to the A gland zone separated by less than 10mm from the proventriculus (B glands). 6. B glands contain the same mucosubstances as in the A glands but no strong acid mucopolysaccharide. 7. The B glands also contain pepsinogen granules in the basal portion of the glands. 8. The A and B glands belong to the same category, however, the A glands are slightly more undifferentiated than the B glands. Namely, a few of the parietal cells and a large number of the strongly PAS- and AB (pH 2.5)-positive cells in the A glands are a symbol of immaturity. 9. There is a mixed zone in both gland regions. 10. The fundic glands have a large number of parietal cells like the B glands. 11. The pepsinogen granules in the fundic glands surpass the B glands in number and size. 12. The fundic glands have no weak and neutral mucopolysaccharides in the glandular lower portion which are contained in the lower portion of the B glands. The lack of the mucosubstances is an indication of the maturity of the glandular cells. 13. Based on the above findings, the B and fundic glands are of the same type, though, the latter is mature and the former is in an immature stage. 14. The so-called cardiac glands are A and B glands. Accordingly, they are not special kinds of glands but undifferentiated fundic glands.
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